:root {
  --bg:#f6f7f4; --paper:#fff; --ink:#182126; --muted:#5c676d; --soft:#7a858a;
  --line:#d8dedb; --accent:#21656c; --accent-deep:#174a50; --accent-pale:#e7f0ef;
  --warm:#b97842; --warm-pale:#f4eee8; --sans:Inter,"Helvetica Neue",Arial,sans-serif;
  --serif:"Iowan Old Style","Palatino Linotype",Palatino,Georgia,serif; --width:68rem; --read:46rem;
}
*{box-sizing:border-box} html{scroll-behavior:smooth} body{margin:0;background:var(--bg);color:var(--ink);font-family:var(--sans);font-size:16px;line-height:1.68;-webkit-font-smoothing:antialiased}
a{color:var(--accent-deep);text-underline-offset:.2em} a:hover{color:var(--accent)} a:focus-visible,button:focus-visible,input:focus-visible,select:focus-visible,textarea:focus-visible{outline:3px solid rgba(33,101,108,.22);outline-offset:3px}
.skip-link{position:fixed;left:1rem;top:-5rem;z-index:100;padding:.65rem .9rem;background:var(--ink);color:#fff;text-decoration:none}.skip-link:focus{top:1rem}
.site-header{position:sticky;top:0;z-index:20;background:rgba(246,247,244,.95);border-top:3px solid var(--accent);border-bottom:1px solid var(--line);backdrop-filter:blur(10px)}
.header-inner{max-width:var(--width);margin:auto;padding:.9rem 1.5rem;display:flex;align-items:center;justify-content:space-between;gap:2rem}.header-actions{display:flex;align-items:center;gap:.8rem;min-width:0}.identity{display:grid;color:var(--ink);line-height:1.2;text-decoration:none}.identity-name{font-weight:720;letter-spacing:-.015em}.identity-role{margin-top:.2rem;color:var(--muted);font-size:.75rem}.site-nav{display:flex;gap:.1rem;flex-wrap:wrap}.site-nav a{padding:.43rem .58rem;color:var(--muted);font-size:.82rem;font-weight:650;text-decoration:none;border-bottom:2px solid transparent}.site-nav a:hover,.site-nav a[aria-current=page]{color:var(--accent-deep);border-bottom-color:var(--accent)}.language-switcher{display:flex;align-items:center;gap:.3rem;flex:0 0 auto;padding-left:.75rem;border-left:1px solid var(--line);color:var(--soft);font-size:.7rem;font-weight:760;letter-spacing:.08em}.language-switcher a{color:var(--muted);text-decoration:none}.language-switcher a:hover{color:var(--accent-deep)}.language-switcher [aria-current=true]{color:var(--ink)}
main{max-width:var(--width);margin:auto;padding:0 1.5rem}.eyebrow,.mini-label{margin:0 0 .8rem;color:var(--accent);font-size:.72rem;font-weight:760;letter-spacing:.12em;text-transform:uppercase}.mini-label{margin-bottom:.55rem;font-size:.67rem}.lead{max-width:var(--read);color:#354148;font-family:var(--serif);font-size:clamp(1.16rem,2vw,1.42rem);line-height:1.5}h1,h2,h3{margin-top:0;line-height:1.18}h1{max-width:50rem;margin-bottom:1.35rem;font-family:var(--serif);font-size:clamp(2.6rem,5.4vw,4rem);font-weight:520;letter-spacing:-.04em}h2{font-size:clamp(1.45rem,2.5vw,2.05rem);letter-spacing:-.025em}h3{font-size:1.08rem;letter-spacing:-.012em}p{margin:0 0 1rem}.text-link{display:inline-block;margin-top:.5rem;font-size:.86rem;font-weight:720;text-decoration-thickness:1px}.button{display:inline-flex;min-height:2.7rem;align-items:center;justify-content:center;padding:.62rem .95rem;border:1px solid var(--accent-deep);border-radius:2px;font-size:.82rem;font-weight:720;text-decoration:none}.button.primary{background:var(--accent-deep);color:#fff}.button.primary:hover{background:var(--accent);color:#fff}.button.secondary{background:transparent;color:var(--accent-deep)}.button.secondary:hover{background:var(--accent-pale)}.hero-actions{display:flex;flex-wrap:wrap;gap:.7rem;margin-top:1.8rem}
.home-hero{display:grid;grid-template-columns:minmax(0,1fr) minmax(17rem,18.5rem);gap:4.25rem;align-items:center;padding:5.5rem 0 4.8rem;border-bottom:1px solid var(--line)}.hero-copy{max-width:49rem}.hero-profile{max-width:18.5rem;justify-self:end;background:rgba(255,255,255,.74);border:1px solid var(--line);border-top:3px solid var(--accent);padding:1.25rem 1.3rem 0;box-shadow:0 12px 35px rgba(24,33,38,.035)}.current-roles p{color:var(--muted);font-size:.82rem;line-height:1.5}.current-roles p:first-child{margin-top:0}.current-roles strong{color:var(--ink)}.current-roles .location{margin-bottom:0;color:var(--soft);font-size:.76rem}.profile-portrait{display:flex;justify-content:flex-end;align-items:flex-end;margin-top:1.05rem;padding-top:.85rem;border-top:1px solid var(--line);min-height:7rem;overflow:hidden}.portrait{display:block;width:5.45rem;height:6.85rem;margin:0 .15rem 0 0;object-fit:cover;object-position:50% 14%;border-radius:2px 2px 0 0}
.page-hero{padding:5rem 0 4.2rem;border-bottom:1px solid var(--line)}.simple-hero>p:not(.eyebrow),.publication-hero>p:not(.eyebrow){max-width:var(--read)}.research-hero{display:grid;grid-template-columns:minmax(0,1fr) 11rem;gap:4rem;align-items:center}.microscopy-figure{width:11rem;margin:0;justify-self:end}.microscopy-figure img{display:block;width:100%;height:auto;object-fit:contain}.microscopy-figure figcaption{margin-top:.6rem;color:var(--soft);font-size:.68rem;line-height:1.4}.compact-hero{padding-bottom:3.5rem}.contact-hero{padding-bottom:2.5rem}.success-page{min-height:58vh}
.editorial-section{padding:4.8rem 0;border-bottom:1px solid var(--line)}.numbered-section,.featured-section{display:grid;grid-template-columns:10rem minmax(0,1fr);gap:3rem}.section-kicker{display:flex;align-items:baseline;gap:.75rem;color:var(--soft)}.section-kicker span{color:var(--warm);font-size:.72rem;font-weight:760;letter-spacing:.08em}.section-kicker p{margin:0;font-size:.72rem;font-weight:720;letter-spacing:.08em;text-transform:uppercase}.reading-column{max-width:var(--read)}.wide-reading{max-width:54rem}.reading-column>p{color:var(--muted)}.reading-column h2{max-width:43rem}.central-question{margin-top:2rem!important;padding:1.2rem 1.35rem;border-left:3px solid var(--warm);background:var(--warm-pale);color:var(--ink)!important;font-family:var(--serif);font-size:1.13rem}.central-question span{display:block;margin-bottom:.35rem;color:var(--warm);font-family:var(--sans);font-size:.67rem;font-weight:760;letter-spacing:.1em;text-transform:uppercase}
.domains-section,.activity-section,.profiles-section{display:grid;grid-template-columns:10rem minmax(0,1fr);gap:3rem}.domain-columns{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:3.5rem}.domain-block{padding-top:1rem;border-top:2px solid var(--accent)}.domain-block p{color:var(--muted)}.activity-list{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:2rem}.activity-list article{padding-top:1rem;border-top:1px solid var(--line)}.activity-list p{color:var(--muted);font-size:.92rem}.section-link{grid-column:2;margin-top:1.5rem}.profile-row{display:grid;grid-template-columns:repeat(4,minmax(0,1fr));gap:1rem}.profile-row a{padding:1rem 0;border-top:2px solid var(--accent);color:var(--ink);text-decoration:none}.profile-row span{display:block;color:var(--soft);font-size:.68rem;letter-spacing:.08em;text-transform:uppercase}.profile-row strong{font-size:.82rem}
.question-list{max-width:54rem}.question-list article{padding:0 0 2.1rem;margin-bottom:2.1rem;border-bottom:1px solid var(--line)}.question-list article:last-child{margin-bottom:0;padding-bottom:0;border-bottom:0}.question-list h2{font-size:1.48rem}.question-list p{max-width:47rem;margin-bottom:0;color:var(--muted)}
.selected-work{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:0 2.2rem;margin-top:2.8rem;border-top:1px solid var(--line)}.selected-work article{padding:1.55rem 0;border-bottom:1px solid var(--line)}.selected-work h3,.selected-work h2{margin-bottom:.65rem;font-size:1.05rem}.selected-work h3 a,.selected-work h2 a{color:var(--ink);text-decoration-thickness:1px}.selected-work p:last-child{margin-bottom:0;color:var(--muted);font-size:.9rem}.section-cta{margin-top:2rem}
.focus-list{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:0 3rem;max-width:56rem}.focus-list article{padding:1.4rem 0 1.6rem;border-top:1px solid var(--line)}.focus-list h2{margin-bottom:.6rem;font-size:1.18rem}.focus-list p{margin-bottom:0;color:var(--muted);font-size:.92rem}.resource-links{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:0 2.5rem;max-width:56rem}.resource-links a{position:relative;padding:1.4rem 2rem 1.6rem 0;border-top:1px solid var(--line);color:var(--ink);text-decoration:none}.resource-links a:after{content:'↗';position:absolute;top:1.25rem;right:.1rem;color:var(--accent)}.resource-links span{display:block;margin-bottom:.45rem;color:var(--accent);font-size:.67rem;font-weight:760;letter-spacing:.08em;text-transform:uppercase}.resource-links h2{margin-bottom:.55rem;font-size:1.15rem}.resource-links p{margin-bottom:0;color:var(--muted);font-size:.9rem}.quiet-notice{max-width:52rem;margin:4.5rem 0 5.5rem 13rem;padding:1.6rem 0;border-top:2px solid var(--warm);border-bottom:1px solid var(--line)}.quiet-notice h2{font-size:1.25rem}.quiet-notice p{color:var(--muted);font-size:.92rem}.quiet-cta{display:flex;align-items:center;justify-content:space-between;gap:2rem;margin:4.5rem 0 5.5rem;padding:1.4rem 0;border-top:1px solid var(--line);border-bottom:1px solid var(--line)}.quiet-cta p{margin:0;font-weight:650}
.featured-section{padding-bottom:4rem}.publication-features{margin-top:0}.record-heading{display:grid;grid-template-columns:10rem minmax(0,1fr);gap:3rem;padding-bottom:1.5rem;border-bottom:0}.publication-list{max-width:58rem;margin-left:13rem;padding:0 0 5.5rem}.strand{margin:3.4rem 0 1.2rem;padding-top:1rem;border-top:1px solid var(--line);color:var(--accent);font-size:.72rem;font-weight:760;letter-spacing:.1em;text-transform:uppercase;scroll-margin-top:7rem}.strand:first-child{margin-top:0;padding-top:0;border-top:0}ol.pubs{list-style:none;margin:0;padding:0;counter-reset:publication}ol.pubs li{position:relative;padding:0 0 1.1rem 2rem;margin-bottom:1.1rem;border-bottom:1px solid rgba(216,222,219,.75);counter-increment:publication}ol.pubs li::before{content:counter(publication);position:absolute;left:0;top:.08rem;color:var(--soft);font-size:.72rem}ol.pubs .venue{display:block;margin-top:.3rem;color:var(--muted);font-size:.8rem}ol.pubs .venue em{font-style:normal}
.contact-layout{display:grid;grid-template-columns:minmax(0,1.45fr) minmax(15rem,.72fr);gap:4rem;align-items:start;padding:2.8rem 0 5.5rem}.contact-form{display:grid;gap:1.15rem;max-width:45rem}.form-field{display:grid;gap:.42rem}.form-field label,.form-check label{font-size:.85rem;font-weight:680}.form-field input,.form-field select,.form-field textarea{width:100%;padding:.78rem .85rem;border:1px solid #aeb8b5;border-radius:2px;background:#fff;color:var(--ink);font:inherit}.form-field textarea{min-height:10rem;resize:vertical}.form-check{display:grid;grid-template-columns:auto 1fr;gap:.65rem;align-items:start;padding:1rem 0;border-top:1px solid var(--line);border-bottom:1px solid var(--line)}.form-check input{width:1rem;height:1rem;margin-top:.25rem}.form-submit{width:max-content;cursor:pointer}.form-note{margin-bottom:0;color:var(--muted);font-size:.78rem}.form-honeypot{position:absolute!important;overflow:hidden!important;clip:rect(0 0 0 0)!important;clip-path:inset(50%)!important;width:1px!important;height:1px!important;margin:-1px!important;white-space:nowrap!important}.contact-side{display:grid;gap:2.5rem;padding-top:.2rem}.plain-links{list-style:none;margin:0;padding:0;border-top:1px solid var(--line)}.plain-links li{padding:.65rem 0;border-bottom:1px solid var(--line)}.plain-links a{font-size:.9rem}.contact-warning{padding-top:1.2rem;border-top:2px solid var(--warm)}.contact-warning h2{font-size:1.1rem}.contact-warning p{color:var(--muted);font-size:.86rem}.legal-content{max-width:var(--read);padding:4rem 0 6rem}.legal-content h2{margin-top:2.4rem;font-size:1.3rem}.legal-content h2:first-child{margin-top:0}.legal-content p{color:var(--muted)}
.site-footer{max-width:var(--width);margin:auto;padding:2.3rem 1.5rem 3.2rem;border-top:1px solid var(--line);display:grid;grid-template-columns:1.25fr 1fr auto;gap:2rem;color:var(--muted);font-size:.78rem}.footer-identity{display:grid}.footer-identity span{margin-top:.15rem}.footer-links{display:flex;flex-wrap:wrap;align-content:start;gap:.35rem 1rem}.footer-links a{color:var(--muted)}.copyright{text-align:right}
@media(max-width:58rem){.site-header{position:static}.header-inner{display:block}.header-actions{margin-top:.75rem}.site-nav{margin-left:-.55rem;min-width:0}.home-hero{grid-template-columns:minmax(0,1fr) 16.5rem;gap:3rem}.hero-profile{max-width:16.5rem}.numbered-section,.featured-section,.domains-section,.activity-section,.profiles-section,.record-heading{grid-template-columns:6rem minmax(0,1fr);gap:2rem}.publication-list{margin-left:8rem}.profile-row{grid-template-columns:repeat(2,minmax(0,1fr))}.quiet-notice{margin-left:8rem}.research-hero{grid-template-columns:minmax(0,1fr) 9rem;gap:2.5rem}.microscopy-figure{width:9rem}.contact-layout{gap:2.5rem}.site-footer{grid-template-columns:1fr 1fr}.copyright{text-align:left}}
@media(max-width:42rem){main{padding:0 1rem}.header-inner{padding:.8rem 1rem}.identity-role{display:none}.header-actions{align-items:center;gap:.5rem}.site-nav{overflow-x:auto;flex-wrap:nowrap;padding-bottom:.15rem}.site-nav a{white-space:nowrap}.home-hero,.research-hero{grid-template-columns:1fr;gap:2.4rem;padding:4rem 0 3.5rem}.hero-profile{width:min(100%,20rem);max-width:20rem;justify-self:start}.profile-portrait{min-height:6.3rem}.portrait{width:5rem;height:6.25rem}.page-hero{padding:4rem 0 3.3rem}h1{font-size:clamp(2.45rem,12vw,3.4rem)}.numbered-section,.featured-section,.domains-section,.activity-section,.profiles-section,.record-heading{display:block}.section-kicker{margin-bottom:1.5rem}.domain-columns,.activity-list,.focus-list,.resource-links,.selected-work{grid-template-columns:1fr}.profile-row{grid-template-columns:1fr}.section-link{grid-column:auto}.microscopy-figure{width:10rem;justify-self:start}.publication-list{margin-left:0}.quiet-notice{margin-left:0}.contact-layout{grid-template-columns:1fr;gap:3rem}.quiet-cta{display:block}.quiet-cta .button{margin-top:1rem}.site-footer{grid-template-columns:1fr}.copyright{text-align:left}}
@media(prefers-reduced-motion:reduce){html{scroll-behavior:auto}}
/* Greek navigation labels are longer; keep the desktop header on one line. */
@media(min-width:58.01rem){html[lang="el"] .header-inner{gap:1rem}html[lang="el"] .header-actions{gap:.45rem}html[lang="el"] .site-nav{flex-wrap:nowrap}html[lang="el"] .site-nav a{padding:.43rem .38rem;font-size:.74rem}html[lang="el"] .language-switcher{padding-left:.55rem}}